Species
What the sources say about this taxon, as written.
diagnosis ·
Plazi
Diagnosis. Tomarus gyas is recognized by the following character combination: frontoclypeal tubercles conical; pronotal tubercle acute and prominent ( Fig. 27F ); pronotal fovea rounded (as wide as interocular distance), smaller in female; metasternum with short setae; protibia tridentate and with an additional basal denticle; apex of metatibia with 21–30 spinules; parameres with a long, acute tooth on each side, apical half strongly narrowed, apices triangularly expanded ( Figs. 16F , 19F ). Taxonomic remarks. Tomarus gyas is one of the most problematic species of the genus because of variable taxonomic characters of external morphology and similar shape of the parameres that sometimes make difficult the differentiation with close species, which have been proposed as synonyms by some authors. After the original description by Erichson (1848) , Arrow (1914) described T. amazonicus to include specimens from Brazilian Amazonia. However, Endrödi (1969) synonymized it under T. gyas based on the approximate shape of the parameres and difficulty to delimitate the differences on the shape of the pronotal fovea and protibiae. More recently, Escalona & Joly (2006) described T. discrepans , which was differentiated only by the shape of the parameres, but it was not recognized in later works such as Ratcliffe et al. (2013) . The present phylogenetic analysis and taxonomic revision shows that T. amazonicus is a valid species not close to T. gyas and differing in several characters, such as the shape of the frontal tubercles, setation on the metasternum, stridulatory apparatus, and internal sac of the aedeagus. Tomarus discrepans , although close to T. gyas , is also a different, valid species.
Counts are recordings at or below each taxon, so an empty branch shows as empty before you click it. A dash means the join view holds no column for that rank and cannot roll up to it.
The rows audioBlast holds for this taxon, each matched to the same Catalogue of Life node. Where a source classifies it differently, its own classification is kept.
| Source | Its name | Its rank | Its family | Record |
|---|---|---|---|---|
| Tomarus gyas | Species | Dynastidae | Plazi/Tomarus gyas |
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