Bioacoustics of Indoseges malkangiri

Diagnosis: Males of I. malkangiri differs from I. sushildutta by having thorn-like macrosetae on the ventral side of the distal half of metatarsi I (in I. sushildutta cuspule-like macrosetae present) ( Fig. 2 A,B ); tibia Iwith a row of short macrosetae on the retrolateral side (in I. sushildutta on the ventro-retrolateral side) ( Fig. 2 B , 7 A ) and lacks any apophysis (in I. sushildutta , tibiae Iwith distinct pointed basal apophysis) ( Fig. 2 A–C , 7 A–C ); femur Iwith a cluster of stridulatory macrosetae but without any tubercle-like apophysis (in I. sushildutta , prominent tubercle-like femoral apophysis present covered with a cluster of stridulatory spines on the distal face) ( Fig. 2 A–D , 7 A–C ); leg formula 2143 (in I. sushildutta leg formula in male 1243); palpal tibiae distinctly incrassate and embolus at the base twists 90° along with distal bulb (in I. sushildutta , slightly incrassate and embolus at the base twists 110°) ( Fig. 3 A–D , 8 A–D ). Females of I. malkangiri differs from all other congers by having two prolateral macrosetae on metatarsi II (in all other species one prolateral macroseta), and a distinct internal genitalia: anterior receptaculum hornlike with a small spherical base and pointed apex; ventral lobe with many elongated entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold narrow, curved and strongly sclerotized forming a very conspicuous and well developed median flap; posterior receptaculum pear shaped ( Fig. 5 A–E ). Females also differ from rest of the species except I. satkosia by having three ventral macrosetae on tibiae IV (in all other species one ventral macroseta) and from I. sushildutta having leg formula 1423 (in I. sushildutta leg formula in female 1243).

Description of holotype male (IPU-17-ARA-554) ( Fig. 1–3 ) . Total length 7.50; carapace 3.89 long, 2.58 wide; abdomen 3.61 long, 2.28 wide; carapace covered with long and short brown hairs on caput and posterior margins; clypeus 0.25 high; eyes posterior row straight ( Fig. 1 D ); ocular group 0.27 long, 0.78 wide; eye diameter ALE 0.14, PME 0.14, PLE 0.14; inter-distances PLE-PME 0.13, ALE-ALE 0.28; abdomen ventrally pale greyish mottled with pale dots; chelicerae retrolaterally with two blunt tubercles at base; labium 0.90 long, 0.51 wide; endites 1.41 long, 0.58 wide; sternum 2.25 long, 1.45 wide, anteriorly marked by thick pale band ( Fig. 1 B ); legs metatarsi Iin ventral distal half with a thorn-like macroseta; tibiae Iwith a row of short macrosetae on retrolateral side ( Fig. 2 B ) and lacks any apophysis ( Fig. 2 A–C ); femur Iwith a cluster of stridulatory macrosetae and lacks prominent tuberclelike apophysis ( Fig. 2 A–D ); leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 3.00, 1.12, 2.61, 2.55, 1.31, 10.59; leg II: 3.21, 1.09, 3.86, 2.78, 1.27, 12.21; leg III: 2.36, 0.86, 1.51, 2.14, 0.82, 7.69; leg IV: 2.98, 1.18, 2.56, 2.51, 1.01, 10.24; palp: 1.25, 0.58, 0.95, -, 0.71, 3.49; leg formula 2143; spines or macrosetae: leg I: fep 16, tip 1-1-1-1, r 1-1-1-1, vp 1-1-1-1, vr 1-1-1-1-1-1-1, mtv 2-0-2; leg II: fep 1-1, tip 1-1-1, r 1-1-1-1, vp 0-0-1, vr 1-1-1, mtvp 1-0-1, vr 1-1-1; leg III: fep 1, tip 1-1-1, r 1-0-1, v 2-1-1-2, mtp 1-0-1, r 1-1-1, v 1-1-3; leg IV: fe d 1-1, ti v 1-1-2, mt vp 1-1-1-2, vr 0-0-0, preening comb with four ventro-retrolateral macrosetae on metatarsi IV; palp: no spine; teeth on paired claws of legs: I: 10, II: 8, III: 7, IV: 6; palp brownish-black; patella and tibia distinctly incrassate; cymbium; bulb pear shaped; embolic duct distinct; embolus at the base twists 90° along with distal bulb; embolus long, tubular, tapers and gradually bends dorsally, and spatulated distally with embolic opening on one side ( Fig. 3 A–D ). Remaining characters same as generic character. Variation in males [n=4; measurements: range (mean±standard deviation)]. Total length 6.88–7.50 (7.26±0.28); carapace. 3.32–3.89 (3.63±0.24) long, 2.09–2.58 (2.28±0.22) wide; abdomen 3.51–3.83 (3.63±0.14) long, 2.01–2.31 (2.21±0.14) wide; clypeus 0.18–0.28 (0.23±0.05) high; ocular group 0.25–0.31 (0.29±0.02) long, 0.61–0.78 (0.69±0.07) wide; eye diameter ALE 0.12–0.19 (0.15±0.03), PME 0.12–0.15 (0.14±0.01), PLE 0.12–0.19 (0.15±0.03); inter-distances PLE-PME 0.08–0.15 (0.12±0.03), ALE-ALE 0.25–0.28 (0.26±0.01); labium 0.63–0.90 (0.78±0.13) long, 0.44–0.51 (0.48±0.04) wide; endites 1.16–1.41 (1.29±0.12) long, 0.43–0.58 (0.51±0.07) wide; sternum 2.00–2.25 (2.13±0.11) long, 1.22–1.45 (1.33±0.12) wide; leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.42–3.00 (2.77±0.25), 0.77–1.18 (1.00±0.19), 2.12–2.61 (2.45±0.23), 2.02–2.55 (2.37±0.24), 1.10–1.34 (1.25±0.11), 8.43–10.59 (9.84±0.98); leg II: 2.62–3.21 (2.95±0.25), 1.01–1.19 (1.10±0.07), 2.43–3.86 (2.43±0.62), 2.61–2.88 (2.77±0.11), 1.19–1.34 (1.26±0.07), 9.86–12.21 (11.05±0.98); leg III: 2.02–2.36 (2.21±0.14), 0.83–0.87 (0.85±0.02), 1.24–1.51 (1.41±0.13), 1.80–2.14 (2.00±0.14), 0.81–0.88 (0.83±0.03), 6.70–7.69 (7.30±0.43); leg IV: 2.55–2.98 (2.97±0.01), 1.04–1.25 (1.22±0.05), 2.06–2.56 (2.43±0.19), 2.18–2.62 (2.57±0.08), 0.79–1.01 (0.96±0.08), 8.62–10.24 (9.61±0.72); palp: 1.07–1.25 (1.14±0.08), 0.54–0.58 (0.56±0.02), 0.87–0.95 (0.92±0.03), -, 0.60–0.71 (0.67±0.05), 3.15–3.49 (3.28±0.15). Description of paratype female (IPU-18-ARA-1274) ( Fig. 4 , 5 ) . Total length 10.96; carapace.4.55 long, 2.75 wide; abdomen 6.41 long, 3.62 wide; carapace covered with long and short brown hairs, relatively more on caput; clypeus 0.30 high; eyes posterior row straight ( Fig. 4 D ); ocular group 0.29 long, 0.88 wide; eye diameter ALE 0.18, PME 0.15, PLE 0.18; inter-distances PLE-PME 0.18, ALE-ALE 0.46; abdomen ventrally pale greyish mottled with pale dots; labium 0.77 long, 0.58 wide; endites 1.45 long, 0.67 wide; sternum 2.64 long, 1.66 wide, anteriorly marked by a thick pale band ( Fig. 4 B ); leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.87, 1.23, 2.10, 1.91, 0.99, 9.10; leg II: 2.74, 1.21, 2.14, 1.95, 1.00, 9.04; leg III: 2.17, 1.00, 1.21, 1.54, 0.72, 6.64; leg IV: 2.88, 1.31, 2.23, 1.93, 0.72, 9.08; palp: 1.10, 0.65, 0.59, -, 0.92, 3.26; leg formula 1423; macrosetae: leg I: fep 1, tip 1-1-1, v 2-2-2, mtvp 1-0-1, vr 1-1-1; leg II: fep 1, tip 1-1-1, vp 1-0-1, vr 0-0-1, mtp 1-1-0, v 2-1-1-2; leg III: ti p 1-1, v 1-2, r 1-1, mt p 1-1-1-1, r 1-1-1-1, v 1-1-1-3; leg IV: ti vp 1-1, vr 0-1, mt vp 1-1- 2, vr 0-0-0, preening comb with four ventro-retrolateral macrosetae on metatarsi IV; palp: ti p 1-1, ta p 13; teeth on paired claws of legs: I: 15, II: 13, III: 11, IV: 9; female genitalia distinct; internal genitalia with a horn-like anterior receptaculum having small spherical base and pointed apex; dorsal lobe spherical; ventral lobe with many elongated entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold narrow, curved and strongly sclerotized forming a very conspicuous and well developed median flap; pear shaped posterior receptaculum ( Fig. 5 A–E ). Remaining characters same as generic character. Variation in females [n=5, measurements: range (mean±standard deviation)]. Total length 8.61–12.80 (10.57±1.61); carapace. 3.69–4.55 (4.17±0.31) long, 2.5–2.75 (2.57±0.13) wide; abdomen 4.92–8.58 (6.40±1.43) long, 2.93–3.62 (3.29±0.26) wide; clypeus 0.24–0.40 (0.32±0.08) high; ocular group 0.25–0.34 (0.30±0.03) long, 0.79–0.94 (0.83±0.10) wide; eye diameter ALE 0.14–0.18 (0.15±0.02), PME 0.13–0.18 (0.15±0.02), PLE 0.14–0.18 (0.15±0.02), inter-distances PLE-PME 0.12–0.18 (0.16±0.03), ALE-ALE 0.36–0.46 (0.39±0.04); labium 0.72–0.78 (0.74±0.03) long, 0.54–0.59 (0.57±0.02) wide; endites 1.26–1.45 (1.35±0.08) long, 0.53–0.67 (0.61±0.05) wide; sternum 2.27–2.64 (2.42±0.16) long, 1.44–1.69 (1.55±0.12) wide; leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.56–2.92 (2.75±0.14), 1.08–1.23 (1.15±0.06), 1.94–2.19 (2.08±0.10), 1.79–2.03 (1.94±0.09), 0.97–1.04 (0.99±0.03), 8.85–9.30 (8.92±0.36); leg II: 2.34–2.81 (2.65±0.19), 1.17–1.27 (1.22±0.04), 1.95–2.15 (2.131±0.11), 1.85–2.08 (2.00±0.10), 0.96–1.02 (0.97±0.05), 9.03–9.37 (8.98±0.42); leg III: 1.91–2.22 (2.08±0.12), 0.98–1.02 (0.98±0.06), 1.07–1.22 (1.17±0.06), 1.38–1.54 (1.49±0.07), 0.65–0.73 (0.69±0.03), 6.00–6.72 (6.41±0.30); leg IV: 2.66–2.97 (2.82±0.12), 1.28–1.31 (1.31±0.05), 2.02–2.34 (2.20±0.12), 1.76–1.93 (1.87±0.09), 0.66–0.77 (0.71±0.05), 8.38–9.44 (8.91±0.41); palp: 0.89–1.10 (1.04±0.09), 0.56–0.65 (0.60±0.04), 0.53–0.59 (0.56±0.03), -, 0.78–1.01 (0.90±0.09), 2.76–3.26 (3.09±0.21); teeth on paired claws of legs: I: 14–15, II: 12–13.