Bioacoustics of Ornebius pullus

Remarks. The specimen collected from Sabah exhibits some differences with O. pullus collected from Belait ( Brunei Darussalam ) and Singapore : legs with more whitish marmoration, supra-anal plate whitish when alive. However, we could not identify clear-cut differences in the genitalia, including both the sclerotized median valve and the membranous lateral valves between the specimen from Sabah and those from Brunei Darussalam ( Fig. 5 ). As Tan et al . (2021) have also suggested that there might be a polymorphic species and that more evidence was needed (e.g., DNA or acoustics) to better understand the species boundaries. At the moment, the external morphology and genitalia morphology are not congruent between specimens from Brunei Darussalam , Singapore and Sabah; we also do not have acoustic data for the Brunei population; and no molecular data are available. Furthermore, the specimen from Sabah also shares some similarities (but also with clear differences) in the external morphology of Ornebius minusculus ( Chopard, 1929 ) from Sipura Island (Sumatra) (http://coldb.mnhn.fr/catalognumber/mnhn/eo/ensif4531), although the genitalia for this species has not been examined. As such, without more specimens from Sabah to examine variations, we refrained from describing it as a new species, tentatively identified it as O. pullus and preferred to revise its taxonomy when the chance arises.