Bioacoustics of Ochodaeus (Ochodaeus) meandrus

Description. Holotype male ( Fig. 3a ). Coleoptera : Scarabaeoidea: Ochodaeidae . Length: 7.4 mm . Width: 3.9 mm . Color: Everywhere uniformly dark reddish brown; surface nitid. Head: Surface lacking tubercles, irregularly punctate; punctures mixed, moderately large and setose, separated by 1–2 diameters. Eyes large, globose; lacking ventral projection of canthus. Antennal club large, subequal to eye in size. Mandibles broadly scooped, shortened (shorter than head), apices falcate; left mandible with large triangular, subacute internal tooth; right mandible with complimenting tooth obsolete, second tooth larger, subacute. Frons weakly convex; surface punctate behind frontoclypeal suture. Clypeus flat with disc relatively long (1/4 as long medially as wide), subtrapezoidal; anterior margin beaded, bead roughly circular but wavy ( Fig. 18 ). Mentum flat emarginated by round fovea in apical half ( Fig. 6 ). Pronotum: Form convex. Surface densely punctate, appearing smooth between punctures laterally, disc with slight indication of tiled granules; punctation mixed, predominantly large setose punctures with few smaller glabrous punctures interspersed. Elytra: Form convex, relatively elongate, elytra together longer than wide. Surface with striae moderately impressed, punctate, surface smooth between punctures (except sutural stria with connecting groove); strial punctures large, separated by 2 diameters, lacking setae. Intervals irregularly punctate; punctures small, setose, with tubercle anteriorly. Legs: Protibia tridentate externally, internal side with short, ventrally directed pollex <1/3 length of apical spur ( Fig. 3b ). Profemur with strong apical tooth. Mesofemur and metafemur with isolated subapical tooth not strongly connected to apex. Venter/Abdomen: Metasternum and abdominal segments sparsely punctate; punctures large with long, erect, golden setae. Pygidium roughly punctate. Stridulatory peg short, capitate ( Fig. 12 ). Male genitalia: Internal sac with narrow arrowhead-shaped patch of sclerites between parameres ( Fig. 9 ). Paratype variation. Sexual dimorphism is not readily apparent in this species. The acute, apical tooth on the profemur is not present in females or smaller males, and the subapical tooth on the mesofemur is also obsolete on most female specimens. The subapical tooth on the metafemur may be reduced but is present on all specimens.

Description. Holotype male ( Fig. 3a ). Coleoptera : Scarabaeoidea: Ochodaeidae . Length: 7.4 mm. Width: 3.9 mm. Color: Everywhere uniformly dark reddish brown; surface nitid. Head: Surface lacking tubercles, irregularly punctate; punctures mixed, moderately large and setose, separated by 1–2 diameters. Eyes large, globose; lacking ventral projection of canthus. Antennal club large, subequal to eye in size. Mandibles broadly scooped, shortened (shorter than head), apices falcate; left mandible with large triangular, subacute internal tooth; right mandible with complimenting tooth obsolete, second tooth larger, subacute. Frons weakly convex; surface punctate behind frontoclypeal suture. Clypeus flat with disc relatively long (1/4 as long medially as wide), subtrapezoidal; anterior margin beaded, bead roughly circular but wavy ( Fig. 18 ). Mentum flat emarginated by round fovea in apical half ( Fig. 6 ). Pronotum: Form convex. Surface densely punctate, appearing smooth between punctures laterally, disc with slight indication of tiled granules; punctation mixed, predominantly large setose punctures with few smaller glabrous punctures interspersed. Elytra: Form convex, relatively elongate, elytra together longer than wide. Surface with striae moderately impressed, punctate, surface smooth between punctures (except sutural stria with connecting groove); strial punctures large, separated by 2 diameters, lacking setae. Intervals irregularly punctate; punctures small, setose, with tubercle anteriorly. Legs: Protibia tridentate externally, internal side with short, ventrally directed pollex <1/3 length of apical spur ( Fig. 3b ). Profemur with strong apical tooth. Mesofemur and metafemur with isolated subapical tooth not strongly connected to apex. Venter/Abdomen: Metasternum and abdominal segments sparsely punctate; punctures large with long, erect, golden setae. Pygidium roughly punctate. Stridulatory peg short, capitate ( Fig. 12 ). Male genitalia: Internal sac with narrow arrowhead-shaped patch of sclerites between parameres ( Fig. 9 ). Paratype variation. Sexual dimorphism is not readily apparent in this species. The acute, apical tooth on the profemur is not present in females or smaller males, and the subapical tooth on the mesofemur is also obsolete on most female specimens. The subapical tooth on the metafemur may be reduced but is present on all specimens.