Bioacoustics of Agyneta sheffordiana

Description: Male : Total length 1.35; carapace length 0.60, width 0.41. CEPHALOTHORAX: Carapace light khaki-brown, shiny, finely reticulate; suffused with dark gray along margin, radiating lines; trident mark present. Sternum suffused with dark gray. Clypeus height 2. Chelicerae light khaki-brown with transverse gray band, not excavated; seta-tipped tubercles absent; promargin four teeth, retromargin four denticles. Cheliceral stridulatory organ ~16 strong striae, well spaced. ABDOMEN: Uniformly light gray. LEGS: Light yellow, prolateral and retrolateral surface of palpal femora and leg femora I–III suffused with dark gray; leg I total length: 2.38, leg III total length: 2.16, Tm I: 0.24, Tm IV: absent. GENITALIA: Palpal retrolateral tibial apophysis absent; dorsal tibial apophysis pointed, rugose; one retrolateral trichobothria and one dorsal ( Fig. 503 ). Cymbium triangular; glabrous depression present with keel ( Fig. 503 ); dorsal cymbial tubercle absent; ventral tubercle triangular, rugose; prolateral notch shallow ( Fig. 504 ). Paracymbium apical pocket short, anterior pocket absent, posterior pocket short ( Fig. 503 ). Embolus tip rounded; basally with three small spines; Fickert’s gland situated medially, globulous; ventral lamella transparent, pointed; thumb long going over the embolus proper ( Fig. 505 ). Embolus proper set apically, of equal part ( Fig. 505 ). Anterior terminal apophysis narrow, tip with few protrusions; posterior terminal apophysis reduced, not well sclerotized; lamella characteristica long, with a basal triangular process, a medially dentate process ( Fig. 506 ) and serrated tip (variable) ( Fig. 507 ). Female : Total length 1.64; carapace length 0.62, width 0.42. CEPHALOTHORAX: Same coloration as male. Chelicerae promargin four teeth, retromargin three denticles. Cheliceral stridulatory organ ~15 striae, well spaced. ABDOMEN: Same coloration as male. LEGS: Same coloration as male; palpal claw absent; total length leg I: 2.45, total length leg III: 2.22, Tm I: 0.22, Tm IV: absent. GENITALIA: Epigynum with proximal part of scape wide, sides parallel; epigynal slits small; pit hook depression very shallow ( Fig. 508 ); lateral lobes large; stretcher small; pit deep ( Fig. 509 ). Median part of scape narrow; genital pores situated at base of lateral lobes pockets ( Fig. 510 ). Internal genitalia with one large, rounded receptacula and a small triangular one ( Figs 509, 510 ).

Description. Male (n=5) : Total length: 1.25–1.47; carapace length: 0.56–0.61; carapace width: 0.38– 0.44; carapace finely reticulate, shiny, light khaki-brown (1245M); radiating lines and carapace margin suffused with dark gray (417M), cephalic region ornamented by a dark gray marking forming a trident, with 3 short erect setae along midline. Sternum dark gray (417M). Chelicerae light khaki-brown (1245M), with wide, transverse gray band; promargin with 3–4 teeth, retromargin with 3–4 denticles. Cheliceral stridulatory organ visible, with ~ 16 striae; associated stridulatory pick at base of palpal femur present. Abdomen uniformly colored, light gray (401M) to dark gray (405M) densely covered with decumbent setae; book lung cover light yellow (127M) finely reticulate. Legs light yellow (129M), prolateral and retrolateral surface of palpal femora and leg femora I–III suffused with dark gray; leg formula 4123; tibia I–IV with two dorsal macrosetae; metatarsus I with dorsal trichobothrium (Tm I 0.21–0.24); Tm IV absent. Palpus length: 0.22–0.24. Palpal tibia with one retrolateral, one dorsal trichobotria ( Fig. 1 ); cymbium with dorsal conical protuberance ( Fig. 1 ) and prolateralo-basal big tubercule (BT) ( Fig. 2 ); paracymbium (P) with large apical pocket (Appo), proximal part bearing six setae ( Fig. 1 ); apical section of embolus (E) long and curved, embolus proper (EP) short and concealed ( Figs 3, 4 ); sperm duct (SD) with well developed Fickert’s gland (FG) located in the embolus ( Figs 3, 4 ); lamella characteristica (LC) long, straight, with one spike at mid-point, apex with small denticles ( Figs 1– 3 ), denticles variable ( Fig. 5 ); anterior terminal apohysis (ATA) spine-like, with a few setae apically ( Fig. 3 ). Female (n=5) : Total length: 1.53–1.69; carapace length: 0.57–0.66; carapace width: 0.41–0.44. Overall coloration as in male. Carapace with 5 erect setae along midline. Chelicerae promargin with 4 teeth, retromargin with 3 denticles. Cheliceral stridulatory organ visible, with ~ 15 striae; associated stridulatory pick at base of palpal femur present. Abdomen densely covered with decumbent setae; book lung cover as in male. Leg formula 4123; palpal tibia and tarsus dark gray (417M); palpal tarsal claw present but highly reduced; tibia I– IV with two dorsal macrosetae; tibial macrosetae longer and heavier than males; metatarsus I with dorsal trichobothrium (Tm I 0.22–0.24); Tm IV absent. Epigynum width: 0.15–0.17. Epigynum with proximal part of scape (PPS) large, side parallel, not going over the pit (Pi) ( Fig. 6 ); pit hook depression (PHD) shallow ( Fig. 6 ); spermathecae (S) consisting of two receptacula: one large, rounded and one small and oblong ( Figs 7, 8 ).