Bioacoustics of Glyphidrilus kotatinggi

Remarks. Five described species from areas near Malaysia include G. kukenthali , G. malayanus , G. gatesi , G. singaporensis , and G. vangviengensis . They are different from the new species with respect to spermathecae, genital markings, clitellum and wing locations. Glyphidrilus kotatinggi sp. n. is most similar to G. singaporensis Shen & Yeo, 2005 , described from a locality nearby (only 45 km apart). Referring to the original description of the latter, G. kotatinggi differs in larger body size (115 + –195 + mm vs. 112–142 mm ), higher segment number (221 + –415 + vs. 163–220), and in the absence of unpaired post-wing genital markings. Further possible differences (clitellum covering 11–14 segments and wings covering 6–7 segments vs. clitellum covering 9–14 segments and wings covering 5–6 segments) are inconclusive due to intraspecific variations in both species that lead to character overlap. For a detailed comparison, we re-investigated the holotype of G. singaporensis and we collected new topotypic material. As a result, we confirm the three above-mentioned differences of G. kotatinggi to G. singaporensis . A comparison of G. kotatinggi with G. singaporensis (specified for holotype , paratypes , and topotypic material) is given in Table 1 . Differences of further species to G. kotatinggi sp. n. are as follows (see also Table 1 ): G. kukenthali from Sarawak differs in slightly longer wings in 18, ½18, 19–24, ½24, longer clitellum in ½13–34, hearts in 7–11, and spermathecae in 14–18. G. malayanus from the Malay Peninsula has a bit shorter wings in 17, ¾18, 18–⅓21, ⅓22, 22, a shorter clitellum in 15–25, 26, intestinal origin in 16, and spermathecae in 14/15–16/17. G. gatesi from Johor , Malaysia has wings in 18, ½19, 19–½24, 24, a shorter clitellum, in ½17, 17, 18–25, ½26, 26, intestinal origin in 18, and spermathecae in 15–17. G. vangviengensis reported from the river bank of Song River at Vangvieng, Laos , has longer wings in 24, 25–31, 32, longer clitellum in 19, 20–35, 36, 37, intestinal origin in 16, and spermathecae are absent. G. bisegmentus sp. n. from Air Banun Pandig, Perak , has shorter wings, shorter clitellum, and genital markings and spermathecae are absent. G. peninsularis sp. n. from Sungei Bantang, has longer wings, a slighlty longer clitellum, four pairs of hearts in 8–11, and spermathecae in 14/15–17/18.